Nathan Fillion portrays Adam Mayfair in Season 4 of Desperate Housewives. Canadian-born actor Nathan Fillion gained critical acclaim and a large cult of fans when he starred as Captain Malcolm Reynolds on the Joss Whedon TV series "Firefly" (2002), having previously been a regular on "One Life to Live" (1968) and "Two Guys, a Girl and a Pizza Place" (1998). Fillion had also appeared in small roles in the films Blast from the Past (1999) and Saving Private Ryan (1998). Whedon gave Fillion a chanc